vol. iv, 402; WG 1400-1500, VIII, 1267) and (2) Elizabeth (E), d. of Sir John Grey (d. 1439), who predeceased his father, Reginald (d. 1440), Lord Grey of Ruthin (GEC. Complete Peerage, VI, p. 159; WG 1400-1500, VIII, 1267). 2 Sir William Griffith II (d. 1531) of Penrhyn m, (1) Jane (W), d. of Thomas Stradling (d. 1480) of St. Donat's, co. Glamorgan (WG 1400-1500, VIII, 1268; Rice Merrick, Morganiae Archaiographia ed. B LI James (South Wales Record Society, vol. 1, 1983), p. 163) and (2) Jane (W), d. of John Puleston of Hafod-y-wern, Wrexham, and Bersham, co. Denbigh and widow of Robert ap Maredudd of Glynllifon, co. Caerns (WG 1400-1500, VIII, 1268; IX, 1454). 3 Sir John Puleston (d. 1551) of Caernarvon m. (1) Gaynor (W), d. of Robert ap Meredydd ap Hwlcyn Llwyd of Glynllifon, co. Caernarvon and (2) Jonet (Sioned) (W), d. of Meredydd ap Jevan ap Robert of Dolwyddelan and Gwydir, co. Caernarvon (H of C 1509-1558, III, 160-1). 4 Sir Rhys Griffith (d. 1580) of Penrhyn m. (1) Margaret (W), d. of Morris ap John ap Meredydd of Clenennau, co. Caernarvon (2) Jane (W), d. of Dafydd ap William ap Gruffydd of Cochwillan, co. Caernarvon, and (3) Catherine (W), d. of Peter Mostyn of Talacre, co. Flint (WG 1400-1500, VIII, 1268; H of C 1509-1558, II, 260-1, where his second wife, Jane, is described as widow of Robert Owen, family and residence not identified). Cardiganshire No knighted landowners with lands wholly or mainly in Wales. Carmarthenshire 1 Sir Rhys ap Thomas (d. 1525) of Dinefwr m. (1) Eva (W), d. of Henry ap Gwilym ap Thomas of Cwrt Henry, Llangathen parish, co. Carmarthen (WG 1400-1500, IV, 645; V 731; Griffiths, Principality, p. 265) and (2) Janet (W), d. of Thomas Mathew of Radyr, co. Glamorgan and widow of Thomas Stradling (d. 1480) of St. Donat's, co. Glamorgan (WG 1400-1500, IV, 645; Rice Merrick, Morganiae Archaiographia ed. B LI James (South Wales Record Society, vol. 1, 1983), p. 163). 2 Sir Griffith ap Rhys (d. 1521) of Dinefwr m. Catherine (E), d. of Sir John St John (d. 1525) of Bletsoe, co. Bedford and Fonmon, co. Glamorgan (WG 1400-1500, IV, 645; Visitations of Bedfordshire (Harleian Society Publications No 19, 1884), p. 53.) His widow later m., as her second husband, Sir Piers Edgcumbe (d. 1539) of Mount Edgcumbe, co. Cornwall (W R B Robinson, 'The Landowners of the Englishry of Gower in the Early Sixteenth Century', Bulletin of the Board of Celtic Studies, xxix, part ii (1981), p. 309). 3 Sir Thomas Philipps (d. circa 1520) of Cilsant, Llanwinio parish, and Picton Castle, co. Pembroke, m. Joan (W), d. of Henry Don of Picton (WG 1400-1500, III, 376; T W Newton Dunn, The Genealogies of the Dwnns of South Wales (Salisbury, 1953), p. 11).